Species
Veronica epacridea
Etymology
Veronica: Named after Saint Veronica, who gave Jesus her veil to wipe his brow as he carried the cross through Jerusalem, perhaps because the common name of this plant is 'speedwell'. The name Veronica is often believed to derive from the Latin vera 'truth' and iconica 'image', but it is actually derived from the Macedonian name Berenice which means 'bearer of victory'.
epacridea: Resembling plants in the family Epacridaceae (a Southern heath family now included in Ericaceae)
Common Name(s)
Hebe
Current Conservation Status
2012 - Not Threatened
Conservation status of New Zealand indigenous vascular plants, 2012
The conservation status of all known New Zealand vascular plant taxa at the rank of species and below were reassessed in 2012 using the New Zealand Threat Classification System (NZTCS). This report includes a statistical summary and brief notes on changes since 2009 and replaces all previous NZTCS lists for vascular plants. Authors: Peter J. de Lange, Jeremy R. Rolfe, Paul D. Champion, Shannel P. Courtney, Peter B. Heenan, John W. Barkla, Ewen K. Cameron, David A. Norton and Rodney A. Hitchmough. File size: 792KB
Previous Conservation Status
2009 - Not Threatened
2004 - Not Threatened
Authority
Veronica epacridea Hook.f.
Family
Plantaginaceae
Brief Description
Low growing shrub with erect twigs bearing pairs of small curved reddish green or green thick fleshy rounded leaves that hardly narrow to a broad leaf stalk that touch those of the opposing leaf inhabiting South Island mountains. Dead leaves persist at base of twigs. Leaves to 9mm long.
Flora Category
Vascular - Native
HEBEPA
The
National Vegetation Survey (NVS) Databank is a physical archive and electronic databank containing records of over 94,000 vegetation survey plots - including data from over 19,000 permanent plots. NVS maintains a standard set of species code abbreviations that correspond to standard scientific plant names from the Ngä Tipu o Aotearoa - New Zealand Plants database.
Structural Class
Dicotyledonous Trees & Shrubs
Synonyms
Hebe epacridea (Hook.f.) Cockayne et Allan nom. superf., nom. illeg., Leonohebe epacridea (Hook.f.) Heads, Hebe epacridea (Hook.f.) Andersen
Distribution
South Island mountains, chiefly on or east of the Main Divide, from the Devil Range, North-West Nelson, to the Eyre and Livingstone Mountains, Southland.
Habitat
Open alpine areas on rock debris or scree. Together with Veronica birleyi and Ranunculus grahamii, it grows al the highest altitudes known for any vascular plant in New Zealand (approximately 2900 m a.s.l in the Malte Brun Range, Aoraki/Mt Cook National Park).
Features
Spreading low shrub (sometimes more or less mat-like) to 0.4 m tall. Branches decumbent or ascending, old stems brown; branchlets green or purplish, puberulent to pubescent or glabrous (rarely), hairs bifarious (usually) or uniform; internodes 1-3 (-4.5) mm; leaves not readily abscising, persistent along the stem for some distance. Leaf bud tightly surrounded by recently diverged leaves. Leaves decussate, connate, usually patent to recurved or erect to erecto-patent; lamina broadly oblong or ovate or elliptic, rigid, somewhat concave or flat (plants from Otago or Aoraki/Mount Cook National Park lack thickened margins), (2.5-) 4-8 (-9) x 2.5-5.5 (-7) mm: apex obtuse or subacute; midrib thickened and evident below (usually forming a prominent keel, except on plants without a thickened leaf margin); margin not cartilaginous, conspicuously thickened (the outward manifestation of a rigid intramarginal vein) or not thickened (on plants that lack a marginal vein), commonly ciliate (toward base and, on one plant from Roys Peak, along entire margin) or minutely papillate or glabrous, sometimes tinged red, entire (usually) or minutely crenulate (rarely) or shallowly toothed (seen on one plant from Otago only); upper surface dark to light green, dull, with many stomata, glabrous; lower surface green, hairy toward base (along connate portion). Inflorescences with 2-8 flowers (per spike), terminal and lateral (arranged as spikes in the axils of little-altered leaves, forming a compact terminal flowering head), unbranched, (0.5-) 0.8-2.6 cm (whole flowering head). Bracts opposite and decussate or lowermost pair opposite, then subopposite or alternate above, connate, ovate or deltoid, obtuse or subacute or acuminate, sometimes hairy outside. Flowers hermaphrodite or female (on different plants). Pedicels absent. Calyx 3.4-5.8 mm; lobes oblong or ovate or elliptic or lanceolate, obtuse or subacute or acuminate. Corolla tube glabrous; tube of hermaphrodite flowers 3.8- 4.8 (-5.4) x 1.6-2.2 mm, cylindric and contracted at base, equalling or longer than calyx; tube of female flowers 2.4-4 x 1.3-1.9 mm, cylindric or funnelform, shorter than (only slightly) or equalling calyx; lobes white at anthesis, elliptic or ovate or obovate (narrowly}, obtuse or subacute, suberect to recurved, shorter than corolla tube. Stamen filaments remaining erect, 0.1-1.2 mm (approximately 0.8-1.2 mm for stamens of hermaphrodite flowers, 0.1-0.4 mm for staminodes of female flowers); anthers yellow or pink to purple, 1.2-2.1 mm; sterile anthers of female flowers pink, 0.8-1.1 mm. Ovary sometimes hairy, 0.8-1.4 mm, apex (in septum view) obtuse or slightly emarginate; ovules 8-18 per locule, in 1-2 layers; style 2.5- 6(-7) mm (generally longer in hermaphrodite flowers than in female flowers), rarely hairy (especially toward base); stigma more prominent in female flowers. Capsules subacute, 2.7-4.5 x 1.5-2.6 mm, sometimes hairy, septicidal split extending ¾-way to base or completely to base, loculicidal split extending¼- (mostly) to ½-way to base. Seeds weakly flattened, ellipsoid or ovoid or obovoid, straw-yellow, 0.8-1 (-1.1) x 0.5-0.7 mm, micropylar rim 0.2-0.3 mm.
Flowering
December - February (-April)
Flower Colours
White
Fruiting
December - April (-September)
Threats
Not Threatened
Chromosome No.
2n = 42
Endemic Taxon
Yes
Endemic Genus
No
Endemic Family
No
Life Cycle and Dispersal
Seeds are wind dispersed (Thorsen et al., 2009).
Taxonomic notes
A widespread and somewhat variable species, distinguished from other members of "Connatae" (Bayly & Kellow, 2006) by: its small rigid leaves, which do not narrow towards the base; retained dead leaves along the length of the stem; and bracts and calyx lobes fringed by long hairs. Leaves are usually strongly keeled with a thickened margin, both characters the result of very thick, woody leaf veins. Plants from Aoraki/Mt Cook National Park and Otago often lack these leaf characters, causing frequent confusion in their identification. Variation is discussed in detail by Kellow el al. (2003).
Attribution
Description adapted by M. Ward from Bayly & Kellow (2006).
References and further reading
Bayly, M.J., Kellow, A.V. 2006. An illustrated guide to New Zealand Hebes. Wellington, N.Z.: Te Papa press pg. 114
Kellow, A. V., Bayly, M. J., Mitchell, K. A., Markham, K. R. and Brownsey, P. J. 2003. A taxonomic revision of Hebe informal group "Connatae" (Plantaginaceae), based on morphology and flavonoid chemistry. New Zealand Journal of Botany 41: 613-35.
Thorsen, M. J.; Dickinson, K. J. M.; Seddon, P. J. 2009. Seed dispersal systems in the New Zealand flora. Perspectives in Plant Ecology, Evolution and Systematics 11: 285-309
This page last updated on 13 Nov 2019